Alumni

2022

Adewunmi Adelaja, MD, PhD

2022
2020
Molecular Biology
Alexander Hoffmann
Resident
Internal Medicine-preliminary, Brigham & Women’s Hospital/ Dermatology, Massachusetts General Hospital
Residency Program Graduate Profile 2021-2022 2016-2020

Cody Aros, MD, PhD

2022
2020
Molecular Biology
Steve Smale
Resident
UCLA Medical Center
Residency Program Graduate Profile 2021-2022 2016-2020

Fitz Gerald Diala, MD, PhD

2022
2020
Molecular Biology
Patricia Johnson
Resident
Vanderbilt University Medical Center
Residency Program 2021-2022 Graduate Profile 2016-2020

Jessica Ong, MD, PhD

2022
2020
Molecular Biology
Karen Reue
Resident
MedStar Georgetown University Hospital
Residency Program Graduate Profile PhD Graduation Year 2021-2022 MD Graduation Year 2016-2020

John Thompson, MD, PhD

2022
2020
Caltech Biology
Linda Hsieh-Wilson
Resident
UCLA
Residency Program Graduate Profile PhD Graduation Year 2021-2022 MD Graduation Year 2016-2020

Kevin Murray, MD, PhD

2022
2020
Molecular Biology
David Eisenberg
Resident
Brown University/Rhode Island Hospital
Residency Program Graduate Profile 2021-2022 2016-2020

LeeAnn Li, MD, PhD

2022
2020
Bioengineering
Song Li
Resident
Johns Hopkins/Bayview
Residency Program Graduate Profile 2021-2022 2016-2020

Peter Pellionisz, MD, PhD

2022
2021
Bioengineering
Maie St. John
Resident
UC San Francisco
Residency Program Graduate Profile 2021-2022 2021-2022

Tran Do, MD, PhD

2022
2020
Molecular Biology
Robert Modlin
Resident
University of Michigan Hospitals-Ann Arbor
Residency Program Graduate Profile 2021-2022 2016-2020

Yeon Joo (Ashley) Kim, MD, PhD

2022
2020
Molecular and Medical Pharmacology
Antoni Ribas
Resident
Internal Medicine, University of Pittsburgh Medical Center
Residency Program 2021-2022 Graduate Profile PhD Graduation Year 2016-2020 MD Graduation Year

2021

Aanand Patel, MD, PhD

2021
2019
Molecular Biology
Margot Quinlan
Resident
Laboratory medicine and Pathology, University Of Washington University Hospital
Residency Program Graduate Profile PhD Graduation Year 2021-2022 MD Graduation Year 2016-2020

Alexander Sun, MD, PhD

2021
2019
Caltech/Biology
Brian M. Stoltz
Resident
NYP Hospital, Columbia University MC
Residency Program Graduate Profile 2021-2022 2016-2020

David Weiss, MD, PhD

2021
2019
Molecular Biology
Robert Modlin
Resident, Preliminary Medicine
Cedars-Sinai Medical Center
Residency Program Graduate Profile 2021-2022 2016-2020

Jonathan Lin, MD, PhD

2021
2017
Bioengineering
Dino DiCarlo
Resident
Emergency Medicine, The Mount Sinai Hospital
Residency Program PhD Graduation Year 2021-2022 Graduate Profile MD Graduation Year 2016-2020 Alumni

Julian Landaw, MD, PhD

2021
2019
Biomathematics
Zhilin Qu
Resident
Anesthesiology, Brigham And Women's Hospital
Residency Program Graduate Profile 2021-2022 2016-2020

Katelyn Masiuk, MD, PhD

2021
2019
Molecular and Medical Pharmacology
Donald Kohn
Chief Scientific Officer
ImmunoVec
2021-2022 Graduate Profile 2016-2020

Lynn Yi, MD, PhD

2021
2019
Caltech/Biology
Lior S Pachter
Resident
Psychiatry, Stanford
Residency Program 2021-2022 Graduate Profile 2016-2020

Matiar Jafari, MD, PhD

2021
2019
Caltech/Biology
Richard Andersen
Resident
Neurological Surgery, UCLA Medical Center
Residency Program Graduate Profile 2021-2022 2016-2020

Michael Liu, MD, PhD

2021
2019
Molecular, Cellular, & Integrative Physiology
Zhilin Qu
Resident
Internal Medicine, Stanford
Residency Program Graduate Profile PhD Graduation Year 2021-2022 MD Graduation Year 2016-2020

Parinaz Abiri, MD, PhD

2021
2019
Bioengineering
Tzung Hsiai
Resident
Internal Medicine, Olive View-UCLA
Residency Program Graduate Profile 2021-2022 2016-2020

Pradeep Rajendran, MD, PhD

2021
2017
Molecular, Cellular, & Integrative Physiology
Kalynam Shivkumar
Resident
Internal Medicine, Massachusetts General Hospital
Residency Program Graduate Profile 2021-2022 2016-2020

Richard Morgan, MD, PhD

2021
2019
Molecular and Medical Pharmacology
Donald Kohn
Resident
Ophthalmology, Duke University Medical Center
Residency Program Graduate Profile 2021-2022 2016-2020

Shivani Thaker, MD, PhD

2021
2019
Molecular and Medical Pharmacology
Heather Christofk
Resident
Massachusetts General Hospital
Residency Program 2021-2022 Graduate Profile 2016-2020

Xuan Tran, MD, PhD

2021
2019
Neuroscience
Shafali Jeste
Resident
Preliminary Pediatrics/Neurodevelopmental Disabilities Childrens National Medical Center
Residency Program Graduate Profile 2021-2022 2016-2020

Xuchen Hu, MD, PhD

2021
2019
Molecular Biology
Stephen Young
Resident
School of Medicine UCLA
Residency Program Graduate Profile 2021-2022 2016-2020

2020

Brian Young, MD, PhD

2020
2018
Molecular Biology
James Wohlschlegel
Resident
Internal Medicine, UCLA
Residency Program Graduate Profile PhD Graduation Year MD Graduation Year 2016-2020 2016-2020

David DiTullio, MD, PhD

2020
2018
Neuroscience
S. Thomas Carmichael
Resident
Internal Medicine, NYU
Residency Program Graduate Profile 2016-2020 2016-2020

Ernest Lee, MD, PhD

2020
2018
Bioengineering
Gerard Wong
Resident
Dermatology, UCSF
Residency Program Graduate Profile PhD Graduation Year MD Graduation Year 2016-2020 2016-2020

Jaspreet Sandhu, MD, PhD

2020
2018
Molecular Biology
Peter Tontonoz
Resident
Internal Medicine, Mass Gen
Residency Program Graduate Profile 2016-2020 2016-2020

Jessica Reddy, MD, PhD

2020
2016
MIT/Biology
Richard Young
PGY-2 Resident
General Psychiatry Residency Program, Johns Hopkins Medicine
Residency Program Graduate Profile 2021-2022 2016-2020

2019

Aleksandr Gorin, MD, PhD

2019
2017
Microbiology, Immunology, & Molecular Genetics
Otto Yang
Resident
Internal Medicine, UCLA
Residency Program Graduate Profile 2016-2020 2016-2020

Andrew King, MD, PhD

2019
2017
Molecular Biology
Guoping Fan
Resident
Dermatology, Wayne State University SOM
Residency Program Graduate Profile 2016-2020 2016-2020

Austin McDonald, MD, PhD

2019
2017
Molecular Biology
Luisa Iruela-Arispe
Resident
Emergency Medicine, Indiana University SOM
Residency Program Graduate Profile 2016-2020 2016-2020

Brandon Thomas, MD, PhD

2019
2017
Molecular Biology
Steve Smale
Resident
Psychiatry, University of Washington Affiliated Hospitals
Residency Program Graduate Profile 2016-2020 2016-2020

Cynthia He, MD, PhD

2019
2017
Neuroscience
Carlos Portera-Cailliau
Resident
Psychiatry, UCSF
Residency Program Graduate Profile 2016-2020 2016-2020

Hua Chai, MD, PhD

2019
2017
Neuroscience
Baljit Khakh
Resident
Psychiatry, UCLA Semel Institute of Neuroscience
Residency Program Graduate Profile 2016-2020 2016-2020

Jason Chen, MD, PhD

2019
2017
Bioinformatics
Giovanni Coppola
Resident
Neurosurgery, Brigham and Women's Hospital
Residency Program Graduate Profile 2016-2020 2016-2020

Jesse Zaretsky, MD, PhD

2019
2017
Molecular and Medical Pharmacology
Antoni Ribas
Fellow
Division of Hematology & Oncology, Washington University
Residency Program Graduate Profile 2016-2020 2016-2020

Jin Park, MD, PhD

2019
2017
Caltech/Biology
Michael Elowitz
Special Volunteer
Cutaneous Microbiome and Inflammation Laboratory
Residency Program Graduate Profile 2016-2020 2016-2020

Joseph Antonios, MD, PhD

2019
2016
Neuroscience
Linda Liau
Resident
Neurological Surgery, Yale-New Haven Hospital
Residency Program Graduate Profile 2016-2020 2016-2020

Kevin Leu, MD, PhD

2019
2017
Bioengineering
Benjamin Ellingson
Resident
Diagnostic Radiology, UCSF
Residency Program Graduate Profile 2016-2020 2016-2020

Minori Ohashi, MD, PhD

2019
2017
Molecular, Cell and Developmental Biology
William Lowry
Resident
Medicine, UCLA
Residency Program Graduate Profile 2016-2020 2016-2020

Narsis Attar, MD, PhD

2019
2017
Molecular Biology
Siavash Kurdistani
Resident
Inernal Medicine, Massachusetts General Hospital
Residency Program Graduate Profile 2016-2020 2016-2020

Philip Bulterys, MD, PhD

2019
2017
Molecular Biology
Jeff F. Miller
Resident
Pathology, Stanford University Programs
Residency Program Graduate Profile 2016-2020 2016-2020

Sheel M. Shah, MD, PhD

2019
2016
Caltech/Biology
Long Cai
Resident
Neurosurgery - UCLA
Residency Program Graduate Profile PhD Graduation Year 2016-2020 2016-2020 MD Graduation Year Alumni

Sophie Rengarajan, MD, PhD

2019
2017
Neuroscience
Elissa Hallem
Resident
Stanford Neurology
Residency Program Graduate Profile 2016-2020 2016-2020

Steven Klein, MD, PhD

2019
2017
Human Genetics
Julian Martinez-Agosto
Resident
Pediatrics/Medical Genetics, Children's Hospital Philadephia
Residency Program Graduate Profile 2016-2020 2016-2020

ZeNan Chang, MD, PhD

2019
2017
Molecular Biology
Yvonne Chen
Resident
Internal Medicine, Barnes-Jewish Hospital , Missouri
Residency Program Graduate Profile PhD Graduation Year MD Graduation Year 2016-2020 2016-2020

2018

Anthony Daggett, MD, PhD

2018
2016
Neuroscience
X. William Yang
Resident, Psychiatry
UCSF Weill Institute for Neurosciences
Residency Program Graduate Profile 2016-2020 2016-2020

Claire Faltermeier, MD, PhD

2018
2016
Molecular Biology
Owen Witte
Resident
Department of Surgery, UCLA
Residency Program Graduate Profile 2016-2020 2016-2020

Esther Nie, MD, PhD

2018
2016
Neuroscience
S. Thomas Carmichael
Resident
Neurology, Stanford
Residency Program Graduate Profile 2016-2020 2016-2020

Genaro Villa, MD, PhD

2018
2016
Neuroscience
Paul Mischel
Clinical Fellow in Neurosurgery
Brigham & Women's Hospital, Harvard
Residency Program Graduate Profile 2016-2020 2016-2020

Gwen Owens, MD, PhD

2018
2016
Caltech/Biology
Pamela Bjorkman
Resident
Anesthesiology, Brigham & Women's Hospital
Residency Program Graduate Profile 2016-2020 2016-2020

Rebecca Rojansky, MD, PhD

2018
2016
Caltech/Biology
David Chan
Resident
Pathology, Stanford
Residency Program Graduate Profile 2016-2020 2016-2020

Trevor Shaddox, MD, PhD

2018
2016
Biomathematics
Marc Suchard
Resident
Psychiaty, UCLA Semel Institute of Neuroscience
Residency Program Graduate Profile 2016-2020 2016-2020

Xavier Gaeta, MD, PhD

2018
2016
Molecular, Cell and Developmental Biology
William Lowry
Fellow
Nephrology, Stanford
Residency Program Graduate Profile 2016-2020 2016-2020

2017

Adam Frank, MD, PhD

2017
2015
Neuroscience
Alcino Silva
Assistant Professor of Clinical Psychiatry and the Behavioral Sciences
Keck School of Medicine of USC
Graduate Profile PhD Graduation Year MD Graduation Year 2016-2020 2011-2015 Alumni

Adam Singer, MD, PhD

2017
2015
RAND
Karl Lorenz
Fellow
UCLA Hematology/Oncology
Graduate Profile 2016-2020 2011-2015

Albert Han, MD, PhD

2017
2015
Neuroscience
Bennett Novitch
Resident
Head & Neck Surgery, UCLA
Graduate Profile 2016-2020 2011-2015

Amit Lakhanpal, MD, PhD

2017
2014
Caltech/Biology
Michael Elowitz
Fellow
Rheumatology Fellowship Training at HSS
Graduate Profile 2016-2020 2011-2015

Anna Reichardt, MD, PhD

2017
2015
Microbiology, Immunology, & Molecular Genetics
Genhong Cheng
Anesthesiology Specialist
Brigham & Women's Hospital
Graduate Profile 2016-2020 2011-2015

Arnav Mehta, MD, PhD

2017
2015
Caltech/Biology
David Baltimore
Postdoctoral Researcher; Clinical Fellow in Hematology/Oncology
Broad Institute of MIT and Harvard; Dana-Farber Cancer Institute
Graduate Profile 2016-2020 2011-2015

Jacob Aptekar, MD, PhD

2017
2014
Neuroscience
Mark Frye
Senior Director
Medidata Solutions, Acorn Al
Graduate Profile 2016-2020 2011-2015

Kevin McEvoy, MD, PhD

2017
2014
Neuroscience
Shafali Jeste
Physician-Scientist
unknown
Graduate Profile 2016-2020 2011-2015

Neelroop Parikshak, MD, PhD

2017
2015
Neuroscience
Daniel Geschwind
Physician-scientist
Regeneron Genetics Center
Graduate Profile 2016-2020 2011-2015

Shiruyeh Schokpur, MD, PhD

2017
2015
MMP
Lily Wu
Resident
Internal Medicine, UC San Diego
Graduate Profile 2016-2020 2011-2015

Wesley Kerr, MD, PhD

2017
2015
Biomathematics
Mark Cohen
Epilepsy Fellow
Univeristy of Michigan
Graduate Profile 2016-2020 2011-2015

2016

Amir Ghaffari, MD, PhD

2016
2013
Microbiology, Immunology, & Molecular Genetics
Genhong Chen
GI and Hepatology Fellow
University of Pittsburgh, Department of Medicine
Graduate Profile 2016-2020 2011-2015

Amy Baohan, MD, PhD

2016
2014
Neuroscience
Joshua Trachtenberg
Resident
Neurological Surgery - Massachusetts General Hospital
Graduate Profile PhD Graduation Year 2016-2020 MD Graduation Year 2011-2015 Alumni

Ivan Osokine, MD, PhD

2016
2014
Microbiology, Immunology, & Molecular Genetics
David Brooks
Postdoctoral Scholar, Laboratory Medicine
UCSF
Graduate Profile 2016-2020 2011-2015

Megan So (Inkeles), MD, PhD

2016
2014
Bioinformatics
Matteo Pellegrini
Dermatologist
Kaiser Permanente
Graduate Profile 2016-2020 2011-2015

Olivia Twu (Choi), MD, PhD

2016
2014
Molecular Biology
Patricia Johnson
Dermatology Physician Scientist, Associate Director
The Janssen Pharmaceutical Companies of Johnson & Johnson
Graduate Profile 2016-2020 2011-2015

Onika Noel, MD, PhD

2016
2014
Molecular Biology
Luisa Iruela-Arispe
Resident Physician
Urologic Surgery, UC Davis Medical Center
Graduate Profile 2016-2020 2011-2015

2015

Aaron Alexander Bloch, MD, PhD

2015
2013
Cambridge/NIH Partnership
Bullmore, Ed (University of Cambridge), Giedd, Jay (NIMH)
Director and Assistant professor
Psychiatry - Children's Hospital of Philadelphia and University of Pennsylvania
2011-2015 2011-2015 Alumni

Alissa Minkovsky (Keegan), MD, PhD

2015
2013
Biological Chemistry
Kathrin Plath
Clinical Research/Medical Director/Research-Related
Amgen, Inc.
PhD Graduation Year Graduate Profile MD Graduation Year 2011-2015 2011-2015

Elif Sozmen (Torres), MD, PhD

2015
2013
Neuroscience
S. Thomas Carmichael
Neurologist
UCSF
Graduate Profile 2011-2015 2011-2015

Jimmy Liu Zhao, MD, PhD

2015
2013
Caltech/Biology
David Baltimore
Physician scientist and hematology/oncologist
Memorial Sloan Kettering Cancer Center
Graduate Profile 2011-2015 2011-2015

Kathryn Cross, MD, PhD

2015
2013
Neuroscience
Marco Iacoboni
Assistant Professor-in-Residence, Neurology
UCLA
Graduate Profile 2011-2015 2011-2015

Kelley O’Donnell, MD, PhD

2015
2013
Neuroscience
Alvaro Sagasti
Psychiatrist and Assistant Professor
Psychiatry - NYU
Graduate Profile 2011-2015 2011-2015 Alumni

Kishan Gupta, MD, PhD

2015
2012
Neuroscience
Michael Hasselmo
Ophthalmologist 
Kaiser Downey Medical Center
Graduate Profile 2011-2015 2011-2015

Lisa Kohn, MD, PhD

2015
2013
Cell & Molecular Pathology
Gay Crooks
HS Clinical Instructor, Medicine
Pediatric Allergy and Immunology - UCLA
2011-2015 2011-2015 Alumni

Oscar Silva, MD, PhD

2015
2013
Microbiology, Immunology, & Molecular Genetics
M. Carrie Miceli
Clinical Assistant Professor, Pathology
Stanford Hosptial and Clinics, Lucile Packard Children's Hospital
Graduate Profile 2011-2015 2011-2015

Scott Knowles, MD, PhD

2015
2013
Molecular and Medical Pharmacology
Anna Wu
Associate Medical Director
Seattle Genetics
Graduate Profile 2011-2015 2011-2015

Victoria Ho, MD, PhD

2015
2013
Neuroscience
Kelsey Martin
Neurology Specialist
UCLA
Graduate Profile 2011-2015 2011-2015

2014

Alejandro Meruelo, MD, PhD

2014
2012
Molecular Biology
James Bowie
Assistant Adjunct Professor, Psychiatry
UCSD
Graduate Profile 2011-2015 2011-2015

David Akhavan, MD, PhD

2014
2012
Molecular and Medical Pharmacology
Paul Mischel
Assistant Professor
Radiation Oncology & Cancer Immunology, University of Kansas Medical Center
Graduate Profile 2011-2015 2011-2015

Erik Anderson, MD, PhD

2014
2012
Molecular Biology
Douglas Black
Radiation Oncologist, Instructor
Radiation Oncology, Beth Israel Deaconess Medican Center; Harvard Medical School
Graduate Profile 2011-2015 2011-2015

Jeffrey Rudie, MD, PhD

2014
2012
Neuroscience
Mirella Dapretto
Adjunct Assistant Professor; Neuroradiologist
UC San Diego; Scripps Health
Graduate Profile 2011-2015 2011-2015

John Colby, MD, PhD

2014
2012
Bioengineering
Elizabeth Sowell
Neuroradiologist 
MarinHealth
Graduate Profile 2011-2015 2011-2015

Jonathan Young, MD, PhD

2014
2012
Caltech/Biology
Michael Elowitz
Ophthalmologist
Legacy Good Samaritan Medical Center, Legacy Salmon Creek Medical Center
Graduate Profile PhD Graduation Year MD Graduation Year 2011-2015 2011-2015 Alumni

Jonathan Zuckerman, MD, PhD

2014
2012
Caltech/Engineering
Mark Davis
Chief and Fellowship Program Director, HS Assistant Clinical Professor
Renal Pathology and Surgical Pathology, UCLA
Graduate Profile 2011-2015 2011-2015

Nathan Hageman, MD, PhD

2014
2011
Neuroscience
Arthur W. Toga
Physician, Director of Research Development
Radiology, University of Toledo; The TouchPoint Solution
Graduate Profile 2011-2015 2011-2015

Omid Kohannim, MD, PhD

2014
2012
Bioengineering
Paul Thompson
Diagnostic Radiologist
LAC-Olive View-UCLA Medical Center
Graduate Profile 2011-2015 2011-2015

Ryan Schmidt, MD, PhD

2014
2012
Neuroscience
Kathrin Plath
Assistant Director, Attending Physician, Assistant Professor
Clinical Genomics Laboratory; Keck School of Medicine of USC
Graduate Profile 2011-2015 2011-2015

Su-Yang Liu, MD, PhD

2014
2012
Microbiology, Immunology, & Molecular Genetics
Genhong Chen
Clinical Research Fellow, Principal Scientist/Pathologist
UCSF; Merck
Graduate Profile PhD Graduation Year MD Graduation Year 2011-2015 2011-2015 Alumni

Valerie Arboleda, MD, PhD

2014
2012
Human Genetics
Eric Vilain
Assistant Professor, Pathology and Laboratory Medicine
UCLA
Graduate Profile 2011-2015 2011-2015

2013

Bahram Razani, MD, PhD

2013
2011
Microbiology, Immunology, & Molecular Genetics
Genhong Cheng
Assistant Professor, Dermatology
UCSF
Graduate Profile 2011-2015 2011-2015

Besim Uzgil, MD, PhD

2013
2011
Neuroscience
Kelsey Martin
Pediatric Neurologist/Epileptologist, Fellow
UCSF, Kaiser Permanente
Graduate Profile 2011-2015 2011-2015

Charles Hummel, MD, PhD

2013
2011
Molecular, Cellular, & Integrative Physiology
Ernest Wright
Partner Physician
Southern California Permanente Medical Group
Graduate Profile 2011-2015 2011-2015

David Van Valen, MD, PhD

2013
2011
Caltech/Engineering
Rob Phillips
Assistant Professor of Biology and Biological Engineering
Biology/Biological Engineering, Caltech
Graduate Profile 2011-2015 2011-2015

Evan Nair-Gill, MD, PhD

2013
2011
Molecular and Medical Pharmacology
Owen Witte
Assistant Professor
Center for Genetics of Host Defense, Internal Medicine, UT Southwestern Medical Center
Graduate Profile 2011-2015 2011-2015

Ezra Rosen, MD, PhD

2013
2011
Neuroscience
Daniel Geschwind
Medical Oncologist
Memorial Sloan Kettering Cancer Center
Graduate Profile 2011-2015 2011-2015

Ghiam Yamin, MD, PhD

2013
2011
Neuroscience
David Teplow
Adjunct Clinical Instructor, Radiology
Kaiser Permanente, Stanford Hospital and Clinics
Graduate Profile 2011-2015 2011-2015

John Phillips, MD, PhD

2013
2011
Caltech/Chemistry
Peter B. Dervan
Assistant Adjust Professor(passed away 9/13/21)
Microbiology, Immunology & Molecular Genetics, UCLA
Graduate Profile 2011-2015 2011-2015

Kelly Bolton, MD, PhD

2013
2011
Cambridge/NIH Partnership Genetic Epidemiology
Paul (NIH Partnership) Pharoah
Assistant Professor
Department of Medicine, Oncology Division, Bone Marrow Transplantation & Leukemia, Washington University
Graduate Profile 2011-2015 2011-2015

Margaret Chang, MD, PhD

2013
2011
Cell & Molecular Pathology
Benhur Lee
Attending Physician, Instructor of Pediatrics
Rheumatology Program, Boston Children's Hospital
Graduate Profile PhD Graduation Year MD Graduation Year 2011-2015 2011-2015 Alumni

Vishal Patel, MD, PhD

2013
2011
Bioengineering
Arthur W. Toga
Neuroradiologist
Mayo Clinic, Jacksonville, Florida
Graduate Profile PhD Graduation Year MD Graduation Year 2011-2015 2011-2015 Alumni

2012

Allen Ardestani, MD, PhD

2012
2010
Neuroscience
Joaquin Fuster
Neuroradiologist
West Cancer Center & Research Institiute, Germantown, TN
Graduate Profile 2011-2015 2006-2010

Caroline W. Vargason (Sham), MD, PhD

2012
2010
Cell & Molecular Pathology
BraunJonathanGordonLynn
Oculofacial Plastic & Reconstructive Surgeon
Jensen Eye Associates, Nampa, ID
Graduate Profile 2011-2015 2006-2010

Christopher Crowley, MD, PhD

2012
2010
Molecular Biology
Todd Yeates
Dermatologist
San Diego Family Dermatology
Graduate Profile 2011-2015 2006-2010

David D. Wu, MD, PhD

2012
2010
Caltech/Engineering
Scott E. Frasier
Assistant Professor
Pulmonary/Critical Care Research, University of Chicago
Graduate Profile PhD Graduation Year MD Graduation Year 2011-2015 2006-2010 Alumni

Jessica Rexach, MD, PhD

2012
2010
Neuroscience
Linda Hsieh-Wilson
Assistant Professor of Neurology
UCLA David Geffen School of Medicine
Graduate Profile 2011-2015 2006-2010

Kellen Winden, MD, PhD

2012
2011
Neuroscience
Daniel Geschwind
Instructor in Neurology
Boston Children's Hospital
Graduate Profile 2011-2015 2011-2015

Ophir Vermesh, MD, PhD

2012
2010
Caltech/Chemistry
James R. Heath
Affiliate, Dean's Office Operations - Dean Other Resident in Radiology
Radiology, Stanford University
Graduate Profile 2011-2015 2006-2010

Steven Jonas, MD, PhD

2012
2010
Material Science & Engineering
Bruce Dunn
Clinical Instructor
Pediatrics, UCLA
Graduate Profile 2011-2015 2006-2010

Tovar Fuller, MD, PhD

2012
2010
Human Genetics
Steven Horvath
Assistant Professor, Psychiatry
UCSF
Graduate Profile 2011-2015 2006-2010

2011

Adam Sperling, MD, PhD

2011
2009
Biological Chemistry
Michael Grunstein
Physician; Instructor
Dana-Farber Cancer Institute; Harvard Medical School
Graduate Profile 2011-2015 2006-2010

Brent M. Kious, MD, PhD

2011
2009
Philosophy
Barbara Herman
Assistant Professor of Psychiatry
University of Utah Health Care
Graduate Profile 2011-2015 2006-2010

Carrie Shilyansky (Holmberg), MD, PhD

2011
2009
Neuroscience
Alcino Silva
Postdoctoral Fellow/Housestaff
Stanford University (Deisseroth Lab) / Stanford Hospital & Clinics
Graduate Profile 2011-2015 2006-2010

Mark W. Dodson, MD, PhD

2011
2009
Molecular Biology
Ming Guo
Gynecological Oncologist
University of Utah
Graduate Profile 2011-2015 2006-2010

Matthew J. Frank, MD, PhD

2011
2009
Cell & Molecular Pathology
Michael Teitell
Assistant Professor of Medicine
Division of Hematology/Oncology, Stanford
Graduate Profile PhD Graduation Year MD Graduation Year 2011-2015 2006-2010

Patricio Riquelme, MD, PhD

2011
2009
Columbia University
Fiona Doestsch
Assistant Professor of Medicine
Oregon Health & Science University
Graduate Profile 2011-2015 2006-2010

Ramin Salehi-Rad, MD, PhD

2011
2009
Chemistry
Michael Jung
HS Assistant Clinical Professor, Medicine
UCLA
Graduate Profile 2011-2015 2006-2010

Stacey Maskarinec, MD, PhD

2011
2009
Caltech/Chemistry
David Tirrell
Assistant Professor of Medicine
Duke University School of Medicine
Graduate Profile 2011-2015 2006-2010

2010

Adam Smith, MD, PhD

2010
2008
Chemistry
Kenneth Houk
Radiologist
Radiation Oncology, University of Kentucky and Affiliates, Lexington KY
Graduate Profile 2006-2010 2006-2010

Arash Shahangian, MD, PhD

2010
2008
Microbiology, Immunology, & Molecular Genetics
Genhong Chen
Head and Neck Surgeon
San Jose Medical Center, Kaiser Permanente
Graduate Profile 2006-2010 2006-2010 Alumni

Asha Iyer, MD, PhD

2010
2008
Caltech/Biology
Richard A. Anderson
Neurological Surgeon
Bayshore Medical Center, JFK Medical Center, Riverview Medical Center
Graduate Profile 2006-2010 2006-2010

Jason Tarpley, MD, PhD

2010
2008
Neuroscience
H. Tad Blair
Director of Stroke and Neurovascular Center, Stroke Neurologist, Vascular Neurology/Clinical Lead
Pacific Stroke and Neurovascular Center
Graduate Profile 2006-2010 2006-2010

Jennifer Fulcher, MD, PhD

2010
2008
Cell & Molecular Pathology
Benhur Lee
Assistant Professor
Division of Infectious Diseases, UCLA
Graduate Profile 2006-2010 2006-2010

Jimmy Donkor, MD, PhD

2010
2008
Human Genetics
Karen Reue
Physician
Emergency Medicine, Community Medical Center and Saint Agnes Medical Center, Fresno, CA
Graduate Profile 2006-2010 2006-2010

Julie (Kate Julie) Baicy, MD, PhD

2010
2008
Neuroscience
Edythe London
Obstetrician; Assistant Professor
Obstetrics & Gynecology - Univ. of New York Langome Medical Center
Graduate Profile 2006-2010 2006-2010

Vaibhav A. Medhekar, MD, PhD

2010
2008
Microbiology, Immunology, & Molecular Genetics
Jeff F. Miller
Primary Care Physician
One Medical Group
Graduate Profile 2006-2010 2006-2010

Vikram C. Anand, MD, PhD

2010
2008
Biological Chemistry
Gregory Payne
Director, Pediatric Infectious Diseases Fellowship Program Attending Physician; Assistant Professor of Clinical Pediatrics
Children's Hospital, Los Angeles, CA; the Keck School of Medicine of USC
Graduate Profile 2006-2010 2006-2010

2009

Calvin T. Yang, MD, PhD

2009
2007
Bioengineering
Jack Judy
Psychiatrist
Clarity Mental Health, Pasadena
Graduate Profile 2006-2010 2006-2010

Esther Melamed, MD, PhD

2009
2007
Neuroscience
Arthur Arnold
Assistant Professor
Department of Neurology, Dell Medical School, University of Texas at Austin
Graduate Profile 2006-2010 2006-2010

Nicholas Nickols, MD, PhD

2009
2007
Caltech/Chemistry
Peter Dervan
Associate Professor-in-Residence
Radiation Oncology, UCLA
Graduate Profile 2006-2010 2006-2010

Rajan P. Kulkarni, MD, PhD

2009
2007
Caltech/Biology
Scott E. Fraser
Associate Professor of Dermatology
OHSU Portland, Oregon
Graduate Profile 2006-2010 2006-2010

2008

Arthur Chou, MD, PhD

2008
2006
Neuroscience
Jeffrey M. Bronstein
Neurosurgeon
Arthur Po-Fei Chou MD Inc. (Private Practice)
Graduate Profile 2006-2010 2006-2010

Bede Broome, MD, PhD

2008
2006
Caltech/Biology
Gilles Laurent
Managing Director
Assured Healthcare Partners
Graduate Profile 2006-2010 2006-2010

Eric Zollars, MD, PhD

2008
2005
Caltech/Biology
Stephen L. Mayo
Medical Director - Early Development - Inflammation
Amgen
Graduate Profile 2006-2010 2001-2005

Jennifer Salstrom, MD, PhD

2008
2006
Human Genetics
York Marahrens
Medical Director; Pediatric Hematologist/Oncologist
Parexel International; Children's Hospital Colorado, Aurora, Colorado
Graduate Profile 2006-2010 2006-2010

Nima Gharavi, MD, PhD

2008
2006
Cell & Molecular Pathology
Judith A. Berliner
Dermatologist
Depratment of Dermatology, Cedars Sinai Medical Center
Graduate Profile 2006-2010 2006-2010

Supriya Saha, MD, PhD

2008
2006
Microbiology, Immunology, & Molecular Genetics
Genhong Chen
Assistant Professor
Oncology, Univesrity of Washington School of Medicine
Graduate Profile 2006-2010 2006-2010

2007

Gagik Oganesyan, MD, PhD

2007
2005
Microbiology, Immunology, & Molecular Genetics
Genhong Chen
Dermatologist
Kaiser Permanente Fontana and Ontario Medical Centers
Graduate Profile 2006-2010 2001-2005

Joseph Hernandez, MD, PhD

2007
2005
Pathology
Linda Baum
Clinical Assistant Professor
Pediatric Allergy & Immunology Clinic, Stanford Children's Health
Graduate Profile PhD Graduation Year MD Graduation Year 2006-2010 2001-2005

Matthew D. Solomon, MD, PhD

2007
2005
RAND
Jose Escarce
Cardiologist
Kaiser Permanente East Bay Service
Graduate Profile 2006-2010 2001-2005

Michael Burgess, MD, PhD

2007
2005
Molecular Biology
Charles Sawyers
Vice President, Early Clinical Development, Immuno-Oncology and Cell Therapy; Assistant Clinical Professor, Hematology/Oncology
Bristol Myers Squibb; UCSF
Graduate Profile 2006-2010 2001-2005

Robert Moccia, MD, PhD

2007
2006
Neuroscience
Kelsey Martin
Principal Scientist, Computational Biology
Pfizer
Graduate Profile 2006-2010 2006-2010

Thomas H. Leung, MD, PhD

2007
2005
Caltech/Biology
David Baltimore
Assistant Professor of Dermatology
Perelman School of Medicine, University of Pennsylvania
Graduate Profile 2006-2010 2001-2005

2006

Belinda H. Tan, MD, PhD

2006
2004
Microbiology, Immunology, & Molecular Genetics
Robert L. Modlin
Medical Dermatologist and Dermatopathologist; Co-Founder and Co-CEO
Good Dermatology; People Science
PhD Graduation Year Graduate Profile MD Graduation Year 2006-2010 2001-2005 Alumni

Carrie Marder, MD, PhD

2006
2004
Neuroscience
Dean Buonomano
Neuroradiologist
SimonMed Imaging, Tucson Arizona
Graduate Profile 2006-2010 2001-2005

Christopher E. Woods, MD, PhD

2006
2004
Neuroscience
Julio L. Vergara
Cardiac Electrophysiologist
Palo Alto Medical Foundation
Graduate Profile 2006-2010 2001-2005

David E. Rex, MD, PhD

2006
2005
Neuroscience
Arthur W. Toga
Neuroradiology Specialist
Kaiser Permanente
Graduate Profile 2006-2010 2001-2005

Derek Welsbie, MD, PhD

2006
2006
Molecular Biology
Charles L. Sawyers
Associate Professor of Ophthalmology
Shiley Eye Institute, UC San Diego Health
Graduate Profile 2006-2010 2006-2010

Houman D. Hemmati, MD, PhD

2006
2006
Caltech/Biology
Marianne Bronner-Fraser
Chief Medical Officer; Co-Founder, Chief Scientific Officer
Vyluma; Levation Pharma, Ltd.
Graduate Profile 2006-2010 2006-2010

Jack Phan, MD, PhD

2006
2004
Human Genetics
Karen Reue
Associate Professor
Radiation Oncology, Univ. of Texas M.D. Anderson Cancer Center
Graduate Profile 2006-2010 2001-2005

Jason N. Itri, MD, PhD

2006
2004
Neuroscience
Chris S. Colwell
Radiologist
Nines Radiology and MetisMD
Graduate Profile 2006-2010 2001-2005

Jeffrey E. Gotts, MD, PhD

2006
2004
Neuroscience
Marie-Francoise Chesselet
Assistant Professor, Medicine
Department of Medicine, UCSF
Graduate Profile 2006-2010 2001-2005

Kyle A. Bernheim, MD, PhD

2006
2004
Caltech/Biology
Richard A. Anderson
Psychiatrist (Child & Adolescent)
Vista Del Mar Family Services/Cedars Sinai Medical Center and Trilliam Family Service, Corvallis, Oregon
Graduate Profile 2006-2010 2001-2005

Meghan L. O’Neil, MD, PhD

2006
2004
Neuroscience
William P. Melega
Psychiatrist
Portland Psychotherapy
Graduate Profile PhD Graduation Year MD Graduation Year 2006-2010 2001-2005 Alumni

Mouhsin Shafi, MD, PhD

2006
2006
Neuroscience
Joaquin M. Fuster
Associate Professor of Neurology; Director, Berenson-Allen Center for Noninvaisve Brain Stimulation
Harvard Medical School
Graduate Profile 2006-2010 2006-2010

Rene J. Trabanino, MD, PhD

2006
2004
Caltech/Chemistry
William A. Goddard
Family Medicine Specialist
Northeast Community Clinics, Los Angeles, CA
Graduate Profile 2006-2010 2001-2005

Sagar A. Vaidya, MD, PhD

2006
2003
Microbiology, Immunology, & Molecular Genetics
Genhong Cheng
Vice President Clinical Development
Travere Therapeutics
Graduate Profile 2006-2010 2001-2005

2005

James P. Howard, MD, PhD

2005
2005
Biological Chemistry
Gregory Payne
Associate Professor, Pediatrics
UCSF School of Medicine, Dept. of Pediatrics
Graduate Profile 2001-2005 2001-2005

Marc Suchard, MD, PhD

2005
2002
Biomathematics
Janet Sinsheimer
Professor of Biostatistics, Biomathematics and of Human Genetics
David Geffen School of Medicine at UCLA
Graduate Profile 2001-2005 2001-2005

Peter Tsai, MD, PhD

2005
2003
Molecular Biology
Hong Wu
Associate Professor, Neurology & Neurotherapeutics
UT Southwestern Medical Center
Graduate Profile 2001-2005 2001-2005

Sameer A. Sheth, MD, PhD

2005
2003
Neuroscience
Arthur W. Toga
Associate Professor of Neurosurgery, Cullen Foundation Endowed Chair, McNair Scholar, Vice Chair of Research
Baylor College of Medicine
Graduate Profile 2001-2005 2001-2005

Steven A. Kushner, MD, PhD

2005
2005
Neuroscience
Alcino J. Silva
Professor and Chair
Neurobiological Psychiatry, Erasmus University Medical Center (The Netherlands)
Graduate Profile 2001-2005 2001-2005

2004

Jia-Perng "Jennifer" Wei, MD, PhD

2004
2002
Molecular Biology
Joan S. Valentine
Medical Director
Agendia, Inc.
Graduate Profile 2001-2005 2001-2005

Marc H. Shomer, MD, PhD

2004
2002
Neuroscience
Marie-Francoise Chesselet
Ophthalmologist
Marc H Shomer MD Phd Inc.
Graduate Profile 2001-2005 2001-2005

Michael Lin, MD, PhD

2004
2002
Biological Chemistry
Michael Greenberg
Associate Professor, Neurology & Bioengineering
Stanford University School of Medicine - Research Intensive
Graduate Profile PhD Graduation Year MD Graduation Year 2001-2005 2001-2005

Osei K. Asamoah, MD, PhD

2004
Physiology
Francisco Bezanilla
Emergency Medicine Specialist
Expressmd Urgent Care Pllc
Graduate Profile 2001-2005

Roger C. Lee, MD, PhD

2004
2002
Biological Chemistry
Larry Zipursky
Dermatologist
Palo Alto Medical Foundation
Graduate Profile PhD Graduation Year MD Graduation Year 2001-2005 2001-2005 Alumni

Ryan B. Green, MD, PhD

2004
2002
Molecular Biology
Judith A. Lengyel
Anesthesiologist
John Muir Medical Center
Graduate Profile 2001-2005 2001-2005

Stanley Raymond Golish, MD, PhD

2004
2002
Computer Science
Sanjiv S. Gambhir
Chief Medical Officer
HCA Healthcare
Graduate Profile 2001-2005 2001-2005

Thomas T. Su, MD, PhD

2004
2002
Molecular Biology Interdepartmental Doctoral Program
David J. Rawlings
Dermatologist
Verdugo Dermatology
Graduate Profile PhD Graduation Year MD Graduation Year 2001-2005 2001-2005 Alumni

2003

Ahmed A. Serag, MD, PhD

2003
2002
Molecular Biology
Todd Yates
Anesthesiologist
Garden Grove Hospital Medical Center
Graduate Profile 2001-2005 2001-2005

Armin G. Jegalian, MD, PhD

2003
2001
Molecular Biology
Hong Wu
Senior Hematopathologist
NeoGenomics Laboratories
Graduate Profile PhD Graduation Year MD Graduation Year 2001-2005 2001-2005

Arnold I. Chin, MD, PhD

2003
2002
Molecular Biology
Genhong Chen
Associate Professor
Urology, David Geffen School of Medicine
Graduate Profile 2001-2005 2001-2005

Doris Y. Chih, MD, PhD

2003
2001
Pathology
H. Phillip Koeffler
Chief of Hospitalist Section of Medical Service
VA Loma Linda
Graduate Profile 2001-2005 2001-2005

Guy L. DiSibio, MD, PhD

2003
2001
Molecular Biology Interdepartmental Doctoral Program
Geraldine Weinmaster
Associate Professor of Pathology and Medical Genetics
Pathology and Medical Genetics - California Northstate University, College of Medicine
Graduate Profile 2001-2005 2001-2005

Hajir Dadgostar, MD, PhD

2003
2001
Biological Chemistry
Genhong Chen
Ophthalmologist, Retina Surgeon
Valley Retina Associates Medical Group, P.C.
Graduate Profile 2001-2005 2001-2005

Ho H. Lee, MD, PhD

2003
2001
Molecular Biology
Genhong Chen
Orthopaedic Surgeon
Kaiser Permanente Fontana Medical Center
Graduate Profile PhD Graduation Year MD Graduation Year 2001-2005 2001-2005 Alumni

Lu Quang Le, MD, PhD

2003
2002
MIMG
Owen Witte
Associate Professor; Dermatologist
Univ. of Texas Southwestern Medical School, Dept. of Dermatology; Children's Health
Graduate Profile PhD Graduation Year MD Graduation Year 2001-2005 2001-2005

Michael Zeineh, MD, PhD

2003
2001
Neuroscience
Susan Y. Bookheimer
Associate Professor of Radiology
Stanford University Medical Center
Graduate Profile 2001-2005 2001-2005

Nader Pouratian, MD, PhD

2003
2001
Neuroscience
Arthur W. Toga
Professor and Chair, Department of Neurological Surgery
UT Southwestern Medical Center
Graduate Profile 2001-2005 2001-2005

Robert A. Lee, MD, PhD

2003
2001
Molecular Biology Interdepartmental Doctoral Program
Hong Wu
Assistant Clinical Professor and Director of Dermatopathology
Dermatology - UCSD and Sutter Health
Graduate Profile PhD Graduation Year MD Graduation Year 2001-2005 2001-2005 Alumni

2002

Angela Lopez, MD, PhD

2002
Biological Chemistry
C. Newton
Oncologist
Cedars-Sinai Medical Center
Graduate Profile 2001-2005

Arun Venkatesan, MD, PhD

2002
2000
Molecular Biology
Asim Dasgupta
Professor of Neurology
Johns Hopkins University School of Medicine
Graduate Profile 2001-2005 1996-2000

Hejin P. Hahn, MD, PhD

2002
2000
Molecular Biology
Linda Baum
Pathologist
Virginia Mason Hospital & Seattle Medical Center - Pathalogy
Graduate Profile 2001-2005 1996-2000

Huidy Shu, MD, PhD

2002
1999
Biological Chemistry
Larry Zipursky
Neurologist
Pali Momi Medical Center and Straub Clinic and Hospital in Aiea, Hawaii
Graduate Profile 2001-2005 1996-2000

Lisan S. Peng, MD, PhD

2002
2000
Molecular Biology
Sherri L. Morrison
Dermatologist
Norman Levine MD Dermatology- Tucson AZ
Graduate Profile 2001-2005 1996-2000

Minghsun Liu, MD, PhD

2002
2000
Microbiology, Immunology, & Molecular Genetics
Jeff F. Miller
National Senior Medical Director and Vice President
Woundtech
Graduate Profile PhD Graduation Year MD Graduation Year 2001-2005 1996-2000 Alumni

2001

Albert Cha, MD, PhD

2001
1999
Neuroscience
Carol Newton
Managing Partner
Frazier’s Life Sciences 
Graduate Profile 2001-2005 1996-2000

Brenton T. Tan, MD, PhD

2001
1999
Molecular Biology
Arnold Berk
Clinical Associate Professor
Stanford University School of Medicine, Department of Pathology
Graduate Profile PhD Graduation Year MD Graduation Year 2001-2005 1996-2000

Charles Chiu, MD, PhD

2001
1999
Biomedical Physics
Phoebe Stewart
Professor, Laboratory Medicine
UCSF School of Medicine
Graduate Profile 2001-2005 1996-2000

Michael Makhinson, MD, PhD

2001
1999
Biological Chemistry
Thomas O'Dell
Psychiatrist 
Psychr & Biobehav Sci / Harbor-UCLA Medical Center
Graduate Profile 2001-2005 1996-2000

Yi-Meng Yen, MD, PhD

2001
1999
Biological Chemistry
Reid Johnson
Orthopedic Surgeon, Assistant Professor of Orthopedic Surgery
Harvard Medical School, Boston Children's Hospital
Graduate Profile 2001-2005 1996-2000

Yifah Yaron, MD, PhD

2001
1999
Biological Chemistry
Judith Gasson
Executive Medical Director
Harpoon Therapeutics
Graduate Profile 2001-2005 1996-2000

2000

Agnieszka "Sheila" Niewmierzycki, MD, PhD

2000
1998
Molecular Biology
Steven Clarke
Pathologist
Oakland
Graduate Profile 1996-2000 1996-2000

Albert Kim, MD, PhD

2000
1998
Physiology
Julio Vergara
Chief Medical Officer
Cytel
PhD Graduation Year Graduate Profile MD Graduation Year 1996-2000 1996-2000 Alumni

Andrew D. Thompson, MD, PhD

2000
1998
Molecular Biology
Christopher Denny
Associate Professor
West Virginia University, Department of Pathology
Graduate Profile PhD Graduation Year MD Graduation Year 1996-2000 1996-2000 Alumni

Andrew F. Cannestra, MD, PhD

2000
1998
Neuroscience
Arthur W. Toga
Neurosurgeon
Baptist Medical Center Jacksonville
Graduate Profile 1996-2000 1996-2000

Benjamin T. Crane, MD, PhD

2000
1996
Neuroscience
Joseph Demer
Professor in departments of Otolaryngology, Biomedical Engineering and Neuroscience
University of Rochester Medical Center
Graduate Profile 1996-2000 1996-2000

Noah Craft, MD, PhD

2000
2000
Molecular Biology
Charles Sawyers
Medical Dermatologist and Dermatopathologist; Co-Founder and Co-CEO
Good Dermatology; People Science
Graduate Profile 1996-2000 1996-2000

Thomas H. Farquhar, MD, PhD

2000
1998
Biomedical Physics
Edward Hoffman
Radiologist
Jefferson Radiology
Graduate Profile 1996-2000 1996-2000

Won-Suk Cho, MD, PhD

2000
1998
Biomathematics
Kenneth Lange
Internal Medicine Physician
San Antonio, TX
Graduate Profile 1996-2000 1996-2000

1999

Daniel Cruz, MD, PhD

1999
1997
Molecular Biology
Mitchell Kronenberg
HS Clinical Professor
David Geffen School of Medicine at UCLA, Div. of Cardiology
Graduate Profile 1996-2000 1996-2000

Daniel M. Sigg, MD, PhD

1999
1999
Physiology
Francisco Benazilla
Nuclear Medicine Specialist
Medical Oncology Associates Ps in Spokane, WA
Graduate Profile 1996-2000 1996-2000

Isla Garraway, MD, PhD

1999
1997
Molecular Biology
Stephen Smale
Associate Professor, Director of Research
David Geffen School of Medicine, UCLA, Dept. of Urology
Graduate Profile 1996-2000 1996-2000

1998

Benjamin S. Braun, MD, PhD

1998
1996
Molecular Biology
Christopher Denny
Associate Professor in Residence, Department of Pediatrics, UCSF; Director, Laboratory for Cell Analysis Core and Associate Director for Shared Resources
UCSF Helen Diller Family Comprehensive Cancer Center
Graduate Profile 1996-2000 1996-2000

Jon Hernandez, MD, PhD

1998
1996
Pathology
Vicente Honrubia
Orthopedist
Surgery, Coordinated Health Systems, Bethlehem, PA
Graduate Profile PhD Graduation Year MD Graduation Year 1996-2000 1996-2000 Alumni

Mark P. Eggena, MD, PhD

1998
1996
Molecular Biology
Jonathan Braun
 Chief Medical Ofiicer, Infectious Disease Specialist 
Penobscot Bay Medical Center, Rockport, Maine
Graduate Profile 1996-2000 1996-2000

Rick Fairhurst, MD, PhD

1998
1996
Molecular Biology
Jonathan Braun
Staff Clinician
National Inst of Allergy and Infectious Diseases, NIH, Malaria Genetics Section, Laboraratory of Malaria and Vector Research
Graduate Profile 1996-2000 1996-2000

William P. Stanford, MD, PhD

1998
1996
Physiology
Charles Markham
Chief Medical Officer
Beverly Hills Institute for Precision Medicine
Graduate Profile 1996-2000 1996-2000

1997

Ajay Nirula, MD, PhD

1997
1995
UT Southwestern
Richard Gaynor
Vice President, Immunology
Eli Lilly and Company
Graduate Profile 1996-2000 1991-1995

Andrei Goga, MD, PhD

1997
1995
Molecular Biology
Owen Witte
Professor, Co-Leader of Breast Oncology Program
Dept. of Cell and Tissue Biology, UCSF; UCSF Helen Diller Family Comprehensive Cancer Center
Graduate Profile 1996-2000 1991-1995

Janine Simmons, MD, PhD

1997
1995
Neuroscience
C. R. Gallistel
Chief
Individual Behavioral Processes Branch - National Institute of Aging, NIH
Graduate Profile PhD Graduation Year MD Graduation Year 1996-2000 1991-1995

Janine Simons, MD, PhD

1997
1995
Neuroscience
C. R. Gallistel
Chief, Individual Behavioral Processes Branch
National Institute of Aging, NIH
Graduate Profile 1996-2000 1991-1995

Roy Avalos, MD, PhD

1997
1995
Microbiology, Immunology, & Molecular Genetics
Debi Nayak
Cardiologist
SD Cardiovascular Associates, Scripps Memorial Hospital, La Jolla, CA
Graduate Profile 1996-2000 1991-1995

Thomas M. Guerrero, MD, PhD

1997
1995
Biomedical Physics
Edward I. Hoffman
Professor, Vice Chair, & Assistant Dean
Department of Radiation Oncology, Beaumont Health, Oakland University William Beaumont School of Medicine
Graduate Profile 1996-2000 1991-1995

1996

Doris Vay Kan Quon, MD, PhD

1996
1994
Microbiology, Immunology, & Molecular Genetics
Patricia Johnson
Hematology Specialist
Hemophilia Treatment Center, Orthopaedic Hospital, LA
Graduate Profile 1996-2000 1991-1995

Mehrdad Matloubian, MD, PhD

1996
1994
Microbiology, Immunology, & Molecular Genetics
Rafi Ahmed
Rheumatologist
UCSF School of Medicine, Dept. of Medicine (Rheumatology/Arthritis)
Graduate Profile 1996-2000 1991-1995

Stephen Lessnick, MD, PhD

1996
1994
Molecular Biology
Christopher T. Denny
Director, Physician, Professor of Pediatrics
Nationwide Children's Hospital, Division of Hematology and Oncology; The Ohio State University College of Medicine
Graduate Profile 1996-2000 1991-1995

1995

Benjamin N. Cheyette, MD, PhD

1995
1993
Molecular Biology
S. Lawrence Zipursky
Psychiatrist
Mindful Health Solutions, Burlingame CA
Graduate Profile 1991-1995 1991-1995

Cynthia A. DeMeester, MD, PhD

1995
1993
Molecular Biology Interdepartmental Doctoral Program
A. Jake Lusis
Pediatrician
Pediatrics - Ascension Borgess Neonatology
Graduate Profile PhD Graduation Year MD Graduation Year 1991-1995 1991-1995 Alumni

Olivia Ong, MD, PhD

1995
1995
Molecular and Medical Pharmacology
Bernard Fung
Ophthalmologist - Glaucoma Specialist
Huntington Memorial Hospital and Methodist Hospital of Southern California
Graduate Profile PhD Graduation Year MD Graduation Year Alumni 1991-1995 1991-1995

Steven P. Hamilton, MD, PhD

1995
1993
Biological Chemistry
Leonard Rome
Psychiatrist, Director of Ketamine for Depression Program
Kaiser Permanente, San Francisco
Graduate Profile 1991-1995 1991-1995

1994

Adam J. Savitz, MD, PhD

1994
1994
Molecular Biology
David I. Meyer
Clinical Assistant Professor of Psychiatry
Weill Cornell Medicine 
Graduate Profile 1991-1995 1991-1995

Beatrice G. Jegalian, MD, PhD

1994
1992
Biological Chemistry
Edward De Robertis
Pediatrician
Pediatrics, Vernon Hall, & Cambridge Health Alliance - Cambridge, Massachusetts
Graduate Profile PhD Graduation Year MD Graduation Year 1991-1995 1991-1995 Alumni

Bradley Fletcher, MD, PhD

1994
1992
Molecular Biology
Harvey Herschman
Associate Professor
Univ. of Florida Health, Hematology/Oncology
Graduate Profile 1991-1995 1991-1995

Keith A. Jr. Wharton, MD, PhD

1994
1992
Molecular Biology
Stephen Crews
VP Medical Director
Ultivue, Cambridge, MA
Graduate Profile 1991-1995 1991-1995

1993

Anthony Dobson, MD, PhD

1993
1991
Microbiology, Immunology, & Molecular Genetics
Lawrence Feldman
Department Chair, Fertility Department
Sutter Health, Palo Alto Medical Foundation
Graduate Profile 1991-1995 1991-1995

Eric C. Svensson, MD, PhD

1993
1991
Biological Chemistry
James Paulson
Associate Professor of Medicine / Co-Director, Physician Scientist Development Program
University of Chicago, Dept. of Medicine (Cardiology), Pritzker School of Medicine
Graduate Profile 1991-1995 1991-1995

Hideki Garren, MD, PhD

1993
1991
Molecular Biology
S. Lawrence Zipursky
Chief Medical Officer
Prothena Corporation plc
Graduate Profile 1991-1995 1991-1995

Jonathan E. Berner, MD, PhD

1993
1991
Neuroscience
Charles Woody
Psychiatrist
Woodinville Psychiatric Associates
Graduate Profile 1991-1995 1991-1995

Joseph A. Garcia, MD, PhD

1993
1991
Microbiology, Immunology, & Molecular Genetics
Richard Gaynor
Professor of Medicine
Division of Cardiology, Columbia University Department of Medicine
Graduate Profile 1991-1995 1991-1995

Michael Teitell, MD, PhD

1993
1991
Molecular Biology
Mitchell Kronenberg
Director, UCLA Jonsson Comprehensive Cancer Center & Professor Pathology, Bioengineering, Pediatrics
Departments of Path & Lab Med, UCLA
Graduate Profile 1991-1995 1991-1995

Pirooz Eghtesady, MD, PhD

1993
1992
Molecular Biology
Mitchell Kronenberg
Chief, Pediatric Cardiothoracic Surgery
Washington University School of Medicine in St. Louis / St. Louis Children's Hospital
Graduate Profile 1991-1995 1991-1995

Robert Hershberg, MD, PhD

1993
Salk Institute/Biology
Mitchell Kronenberg
Venture Partner
Frazier Life Sciences
Graduate Profile 1991-1995

Wes Lee, MD, PhD

1993
1991
Molecular Biology
Arnold Berk
Medical Oncologist
Providence Medical Group Santa Rosa - Cancer Center
Graduate Profile PhD Graduation Year MD Graduation Year Alumni 1991-1995 1991-1995

William M. Gillespie, MD, PhD

1993
1993
Molecular Biology
Charles Woody
Psychiatrist
William M Gillespie & Associates
Graduate Profile 1991-1995 1991-1995

1992

Dean W. Felsher, MD, PhD

1992
1991
Molecular Biology
Jonathan Braun
Professor of Medicine (Oncology) and of Pathology
Stanford University School of Medicine
Graduate Profile 1991-1995 1991-1995

Sanjiv S. Gambhir, MD, PhD

1992
1990
Biomathematics
Sung Cheng Huang
Virginia and D.K. Ludwig Professor in Cancer Research/Chairman of the Department of Radiology (passed away 07/18/2020)
Stanford University School of Medicine
PhD Graduation Year Graduate Profile MD Graduation Year 1991-1995 Alumni 1987-1990

Stewart H. Lecker, MD, PhD

1992
1990
Molecular Biology Interdepartmental Doctoral Program
William Wickner
Assistant Professor and Attending Physician
Nephrology - Harvard Medical School and Beth Israel Deaconess Medical Center
Graduate Profile PhD Graduation Year MD Graduation Year Alumni 1991-1995 1987-1990

1991

Barry Paw, MD, PhD

1991
1989
Biological Chemistry
Elizabeth Neufeld
Associate Professor of Medicine (passed away 12/27/2017)
Harvard Medical School Brigham & Women’s Hospital Division of Hematology
PhD Graduation Year Graduate Profile MD Graduation Year 1991-1995 Alumni 1987-1990

David E. Krantz, MD, PhD

1991
1991
Molecular Biology Interdepartmental Doctoral Program
S. Lawrence Zipursky
Professor in Residence
Psychiatry and Biobehavioral Sciences - UCLA
Alumni 1991-1995 1991-1995

Eric Toloza, MD, PhD

1991
1990
Physiology
Jared Diamond
Thoracic and Cardiac Surgeon
Thoracic Oncology - H. Lee Moffitt Cancer Center and Research Institute
Graduate Profile PhD Graduation Year MD Graduation Year Alumni 1991-1995 1987-1990

1989

David H. Rowitch, MD, PhD

1989
1987
University of Cambridge/Biochemistry
Richard Perham (University of Cambridge)
Professor and Head of Pediatrics, Wellcome Trust Senior Investigator (UK) and Adjunct Professor (UCSF)
Department of Pediatrics - University of Cambridge and UCSF
Alumni 1987-1990 1987-1990

Emil Kakkis, MD, PhD

1989
1988
Biological Chemistry
Kathryn Calame
Founder, President, and Chief Executive Officer
Ultragenyx Pharmaceutical Inc.
Graduate Profile PhD Graduation Year MD Graduation Year Alumni 1987-1990 1987-1990

Hal F. Yee, MD, PhD

1989
1990
Physiology
Glen Langer
Chief Medical Office/ Chief Deputy Director
Health Services, Clinical & Medical Affairs - Los Angeles County Department of Health Services
Graduate Profile PhD Graduation Year MD Graduation Year Alumni 1987-1990 1987-1990

Lily Wu, MD, PhD

1989
1989
Molecular Biology
Arnold Berk
Professor, Departments of Molecular and Medical Pharmacology, Urology, and Pediatrics
UCLA
Graduate Profile PhD Graduation Year MD Graduation Year Alumni 1987-1990 1987-1990

1988

Jeff Bronstein, MD, PhD

1988
1988
Neuroscience
Claude Wasterlain
Fred Silton Family Chair in Movement Disorders and Professor
Neurology - UCLA
Alumni 1987-1990 1987-1990

William R. Wilcox, MD, PhD

1988
1989
Molecular Biology Interdepartmental Doctoral Program
David Eisenberg
Professor
Department of Human Genetics - Emory University
PhD Graduation Year Graduate Profile MD Graduation Year Alumni 1987-1990 1987-1990

1987

Hyo-Chun Yoon, MD, PhD

1987
1989
Molecular Biology Interdepartmental Doctoral Program
Richard Dickerson
Adjunct Professor/ Radiologist
Radiology and Imaging Sciences (UoU) - University of Utah and Kaiser Permanente Moanalua Medical Center
Graduate Profile Alumni 1987-1990 1987-1990